13191.tex

\section{Funding}
\label{funding} 
This work was supported by a grant from the David and Lucille Packard Foundation to PJA. Hollis Gear and Nauticam provided grants for equipment used in directed surveys. The funders had no role in study design, data collection and analysis, 
decision to publish, or preparation of the manuscript.

Competing Interests: The authors have declared that no competing interests exist.